As the name implies, Nightly typically receives patches on a day-to-day basis and new features as soon as they become ready. It represents the bleeding edge of Firefox development, so the likelihood of encountering instability, security vulnerabilities, and major bugs is high.
The Nightly or pre-alpha branch is intended for testing and debugging purposes. Some instability and bugs should be expected. The Developer Edition or alpha branch of Firefox replaced the Aurora branch and is geared towards Web developers and includes extra development tools and a special theme. In the course of a typical six week beta run, there are between seven and thirteen builds released. It is generally quite stable with most remaining bugs being specific to certain configurations or drivers. The Beta branch of Firefox is geared towards home users who want to try new features a few weeks early while contributing to Firefox development. 
It receives a major update every four weeks if all goes according to plan, but security and stability patches are issued as needed in the meantime.
The Release branch of Firefox is geared towards the the average home or office Web user. These are Release, Beta, Dev Edition, Nightly, and ESR. Mozilla offers five officially compiled branches or channels of Firefox. Please use Pale Moon for those olde-skool aesthetics. It will no longer receive security or stability updates. Unfortunately, the v3.6.x series of Firefox is now unsupported. 3.3.1 Adblocking, privacy, and security. 3.1 WebExtension support and legacy add-ons.
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